Where the GM2 story
goes next.

Research directions documented through the laboratory and Bose Institute profiles, presented here as a forward-looking research portfolio.

Y

GM2–Hippo–YAP/TAZ

Identify upstream receptors or components linking GM2 to Hippo pathway inactivation, YAP/TAZ nuclear signalling, EMT and metastasis.

CAF

Tumour–Stroma Crosstalk

Define how GM2-driven tumour–stromal interactions and cancer-associated fibroblast signatures contribute to metastatic behaviour.

3D

GM2-Synthase Structure

Biochemical and structural characterization of GM2-synthase using molecular, mutational and future high-resolution structural approaches.

WHAT WE DISCOVERED

Follow the
evidence.

A compact view of the laboratory's scientific trajectory—from gene regulation to the behaviour of the tumour cell and its ecosystem.

DISCOVERY 01 · GENE REGULATION

Why GM2-synthase rises.

Our work has investigated transcriptional and chromatin mechanisms underlying B4GALNT1/GM2-synthase deregulation in cancer.

ChromatinTranscriptionGM2 synthesis
DISCOVERY 02 · SIGNALLING

GM2 becomes a signalling cue.

GM2-associated membrane signalling engages integrin-linked pathways and the FAK/Src-ERK axis, connecting lipid abundance to malignant cell behaviour.

GM2IntegrinFAK/SrcERK/EGR1
DISCOVERY 03 · TUMOUR PROGRESSION

A lipid can change phenotype.

GM2-driven signalling is linked with EMT, migration, invasion and anchorage-independent growth, providing a mechanistic bridge to metastatic competence.

EMTMotilityInvasionMetastasis
RESEARCH MAP · THE GM2 AXIS

One lipid. Multiple scales of biology.

Follow the laboratory's central mechanistic thread from gene regulation to tumour-cell signalling, microenvironmental remodelling and metastatic growth.

B4GALNT1
GOLGI
GM2 ↑
MEMBRANE
+ SIGNAL
FAK / SRC
→ ERK / EGR1
EMT · AIG
· INVASION
SHEDDING
→ TME / CAF
METASTATIC
GROWTH
GM2 biology
Signalling
Tumour phenotype
Tumour microenvironment
SCIENTIFIC BASIS

What is established
and what remains open.

The website distinguishes the laboratory's GM2-specific mechanistic findings from broader evidence on ganglioside-mediated tumour-microenvironment effects, while presenting the overall sequence as a mechanistic research model rather than implying that every TME effect has already been proven to be GM2-specific.

GM2-specific evidence

GM2 has been shown to interact with integrin and modulate downstream FAK/Src/ERK signalling, promoting migration and invasion; recent work identifies MEK–ERK–Egr1-dependent EMT as a mechanistic axis.

Gangliosides in the TME

Tumour-derived gangliosides can be shed into the microenvironment and influence immune and myeloid compartments. GM2 has specifically been reported to inhibit macrophage Fc-receptor expression and to affect dendritic-cell function in experimental systems.

The research opportunity

How GM2-driven tumour-cell signalling connects to stromal, vascular and immune remodelling in a living tumour remains an important mechanistic question—and a natural next chapter for tumour glycobiology.

Core Scientific Directions

From Membrane Lipids to Oncogenic Programmes

Our research connects glycosphingolipid biology with downstream cell signalling, transcription factors, epigenetic regulation, and aggressive cancer-cell phenotypes.

01 Functional Biology

Genetic Manipulation of Gangliosides

Genetic manipulation of selected gangliosides to define their roles in migration, invasion, anchorage-independent growth and tumour progression.

02 Signalling Pathways

Downstream Signalling Cascades

Dissecting how GM2 engages downstream signalling programmes, including MEK/ERK/Egr1 and transcriptional pathways linked to aggressive cancer phenotypes.

03 Epigenetics

Chromatin & Enzyme Regulation

Uncovering epigenetic mechanisms regulating GM2-synthase expression, including the Sp1–HDAC1–p300 regulatory axis.

04 Metastasis Dynamics

EMT & Metastatic Transition

Understanding how glycolipid-driven changes influence cellular migration, epithelial-mesenchymal transition (EMT), and systemic metastatic behaviour.

05 Genome Engineering

Targeted Gene-Editing Tools

Using targeted genetic tools including TALEN-based editing and contemporary gene-regulatory approaches to interrogate causal disease mechanisms.

06 Translational Oncology

Biomarkers & Therapy Response

Exploring whether mechanistic insights into tumour-associated glycolipids can reveal reliable biomarkers and novel therapeutic opportunities.

SCIENTIFIC ENGAGEMENT

From Kolkata to
global collaborations.

Recent activities include invited lectures, scientific chairing, conference participation and the initiation of a Bose Institute–FAIR/GSI collaboration.

DE

March 2026: visit to FAIR-GSI, Darmstadt, including the lecture “Glycobiology in Tumorigenesis: The GM2 Saga” and initiation of collaboration with Bose Institute.

IACR

Invited to chair the “Immunotherapy—Achievements & Future Directions” session at the 2026 annual meeting in Srinagar.

p300

May 2026: invited lecture at BRIC-NIBMG, Kalyani on transcriptional control of the GM2-synthase gene in cancer.

03 · THE GM2 STORY

A scientific journey
built around one
lipid.

Selected milestones from the laboratory’s published work, assembled from the existing Tumour Glycobiology Laboratory website.

2015

GM2/GD2-synthase

Targeted editing linked ganglioside synthesis with anchorage-independent growth and anoikis resistance.

2016

Migration

GM2 was shown to mediate tumour-cell migration through integrin-associated signalling.

2019

Epigenetics

Histone H3 acetylation and loss of the Sp1–HDAC1 complex were linked to GM2-synthase de-repression in RCC.

2026

EMT programme

GM2 was reported to induce EMT through a MEK/ERK/Egr1-dependent transcriptional programme.

2026

p300 mechanism

dCas9-targeted proteome profiling revealed p300-mediated reciprocal SMAD/SP1 regulation driving GM2-synthase transcription.

What the animation represents: GM2 is shown as a membrane-associated signalling cue that can influence cancer-cell behaviour through Integrin/FAK/Src and MEK–ERK–Egr1 pathways, while gangliosides released into the extracellular milieu can modulate immune and myeloid components of the tumour microenvironment. The latter is a broader ganglioside effect; the specific contribution of GM2 to TME remodelling remains an active research question.

GM2
RESEARCH SIGNATURE

From a tumour-derived glycolipid to a regulatory network.

The laboratory's recent work connects GM2 abundance to signalling and transcriptional programmes that help explain cancer-cell plasticity and progression.

GM2 Glycolipid focus
MEK/ERK Signalling axis
Egr1 Transcriptional effector
p300 Regulatory hub
THE GM2 EFFECT

Why is GM2 high?
What does GM2 do?

plasma membrane → signalling → malignant phenotype → shedding → CAF/TME → metastatic growth. transcriptional deregulation of the GM2-synthase gene increases GM2 production; GM2 accumulates on the tumour-cell membrane, drives pro-tumorigenic signalling and is actively shed into the tumour microenvironment, where gangliosides can modulate immune, myeloid and stromal compartments.

GM2 biosynthesis: ganglioside synthesis proceeds through the secretory pathway: glucosylceramide → lactosylceramide → GM3, followed by Golgi-localized B4GALNT1/GM2-synthase adding GalNAc to GM3 to form GM2; gangliosides are then trafficked to the plasma membrane. The animation is a schematic of this pathway, not a quantitative metabolic map.

01

In cancer, deregulation of the B4GALNT1 gene encoding GM2-synthase increases GM2-synthase transcription. Our work links this de-repression to altered chromatin regulation, including loss of the Sp1–HDAC1 repressive state; recent work identifies p300-mediated reciprocal regulation of SMAD and SP1 as a driver of GM2-synthase transcription in RCC.

02

Increased GM2-synthase activity generates more GM2, producing an aberrantly GM2-rich tumour-cell membrane. GM2 is not merely a passive marker: it participates in membrane microdomains and receptor-associated signalling.

03

Membrane-associated GM2 can engage signalling machinery including integrin-associated FAK/Src and the MEK–ERK–Egr1 axis, promoting EMT-associated changes, migration and invasion.

04

Tumour cells actively shed gangliosides into the extracellular milieu. Shed gangliosides can influence immune and myeloid compartments and contribute to an immunosuppressive, tumour-supportive microenvironment. The precise GM2-specific contribution to each TME compartment.